DNS is a two-way street.
1. You type "myGreatIBMi.com" into a client and that host is resolved to address 1.2.3.4 and is used to connect. That uses the DNS servers configured on the client. I'd say that in most cases, those will be network DNS servers set by DHCP when your client connects to the network.
2. You have an IBMi app that's attempting to connect to a Google webservice. The app uses a URL that starts with "google.com". Your IBMi needs to have DNS servers defined so it can resolve that host to the IP address.
I imagine that #1 is much more common than #2.
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